Evolutionary implications of stipule occurrence in Permo-Carboniferous wetland forests dominated by Psaroniaceae
Weiming Zhou, Wenjun Sun, Ting Wang, Dandan Li, Josef Pšenička, Christopher Hill, C. Kevin Boyce, Jun Wang
The paper identifies the earliest known stipule structure in the plant kingdom from the early Permian Wuda Tuff Flora, providing insights into the evolutionary origins of stipules in Psaroniaceae.
bioRxiv · Jun 23, 2025 · PreprintPaleontology