Isaac Klapper

The paper models resource mismatch in phototrophic biofilms, predicting a steady-state phototroph/heterotroph ratio, which is supported by field observations.

The paper models water dynamics in subaerial biofilms and their interaction with porous stone, showing how biofilms can draw water from substrates to counteract evaporation.

The paper proposes a kinetics-free methodology combining cellular metabolic flux analysis with macro-scale microbial community models to study biofilm communities.